HINT 20G: Half of the *T* alleles are transmitted randomly through females, and so are of type *T*^{p} with probability *p*. Of the other half, transmitted through males, a fraction (1 – *u*) are transmitted randomly through matings with *P*^{V} females and the remaining half are transmitted via *P*^{U} females and must be of type *T*^{P}.