NOTE 17G: The additive genetic variance can also be calculated by summing the variance contributed by the two loci and summing over the two genomes, as 2((0.65)2 + (0.1)2) = 0.867. The slight difference arises because of slight associations (i.e., linkage disequilibria) between the two loci: The red allele is slightly more likely to be found with the black allele and blue with green, which slightly reduces the variance of breeding value.